(900+ acres plus the 550-acre proposed addition, Section 23)
Features found only in the Proposed RNA Addition (Sec. 23)
Current and Future Research Values:
Research and restoration practices to counteract the regional decline of whitebark pine, a candidate for listing as Threatened or Endangered under the ESA
Studies of climatic changes over the past 600 years, and on-going climate change--Alpine larch and whitebark pine produce growth-ring sequences highly sensitive to climatic fluctuations.
Provides a valuable ecological baseline (reference) area for understanding natural conditions in the absence of human influences
Several investigations involving scientists and graduate students from the Rocky Mountain Research Station, universities, and other institutions have been conducted in recent years (see attached list). Proximity of this RNA to major research institutions makes it a prime area for research.
Vulnerability to Impacts from Human Activities:
1917. The conifers of the Northern Rockies. J. E. Kirkwood. U.S. Dept. of Interior, Bureau of Education, Bulletin No. 53.
1963. Zonal great soil groups in western Montana. T. J. Nimlos. Proc. Montana Academy of Sci. 23:3-13. (describes brown podzolic soil on Carlton Ridge)
1968. Chemotaxonomic investigation of hybridization between Larix occidentalis and Larix lyallii. A preliminary study. G. M. Knudsen. M.S. thesis, University of Montana, Missoula. (This and the following study used samples of alpine larch
1969. Hybridization of western and alpine larch. C. E. Carlson and G. M. Blake. Montana Forest Conservation Experiment Sta. Bull. 37.
1970. Ecology of alpine larch (Larix lyallii Parl.) in the Pacific Northwest. S. F. Arno. Ph. D. disserta., Univ. of Montana.
1972. (same title as above). S. F. Arno and J. R. Habeck. Ecological Monographs 42: 417-450.
1977. Forest habitat types of Montana. R. D. Pfister and others. USDA Forest Service, Intermountain Forest and Range Exp. Sta., Gen. Tech. Rep. 34. (Uses data from alpine larch communities on Carlton Ridge to characterize the
1987. Establishment record for Carlton Ridge Research Natural Area, within the Lolo National Forest, Missoula County, Montana. J. R. Habeck. (signed by Forest Service Deputy Chief George Leonard. Unpublished report on file at the Lolo
1988. Research natural areas in the Northern Region: A guidebook for scientists and educators. J. R. Habeck. USDA Forest Service, Northern Region/IntermountainResearch Station. Review Draft. (at www.rna.nris.state.mt.us/rna) (provides a synopsis of physical and ecological characteristics of the Carlton Ridge RNA including a map.)
1990. Hybrid larch of the Carlton Ridge Research Natural Area . . . C. E. Carlson and others. Natural Areas Journal 10(3):134-139. (describes the RNA as a whole, possible expansion of it, and quantifies the characteristics of alpine larch, western larch, and hybrid larch in the rocky outcrop community)
1991. Foliar terpenes of a putative hybrid swarm (Larix occidentalis x Larix lyallii) in western Montana. C. E. Carlson and others. Canadian Jour. of Forestry Research 21:876-881. (This study used samples from alpine larch on the upper slopes in the Carlton Ridge RNA and apparent hybrids from the rocky outcrop site to assess the nature of larch hybridization. Complex introgressive hybridization is indicated by
1993. Cone and seed morphology of western larch (Larix occidentalis), alpine larch (Larix lyallii), and their hybrids. C. E. Carlson and L. J. Theroux. Canadian Jour. of Forestry Research 23:1264-1269. (The Carlton Ridge RNA rocky outcrop site was used to compare and contrast cone and seed morphology among the two larch species and hybrids.)
1994. Germination and early growth of western larch (Larix occidentalis), alpine larch (Larix lyallii), and their reciprocal hybrids. C. E. Carlson. Canadian Jour. of Forestry Research 24:911-916. (Alpine larch trees from the upper slopes of Carlton Ridge RNA were used in experimental cross-pollinations with western larch from another area. Hybrid
1997. Botanical reconnaissance of Carlton Ridge Research Natural Area: Mid- to high-elevation subalpine habitats. C. M. Stalling. USDA Forest Service, Intermountain Research Sta., Research Note 431. (This publication provides an ecological characterization of the area and a complete inventory of its vascular plant flora, consisting of 77 species.)
2003. Fire-climate-vegetation interactions in subalpine forests of the Selway-Bitterroot Wilderness Area, Idaho and Montana, USA. K. F. Kipfmueller. Ph.D. Dissertation, University of Arizona, Tucson, AZ.
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